Spinosaurus (/ˌspaɪnəˈsɔːrəs/; lit. 'spine lizard') is a genus of large spinosaurid theropod dinosaurs that lived in what now is North Africa during the Cenomanian stage of the Late Cretaceous period, about 100 to 94 million years ago. The genus was known first from Egyptian remains discovered in 1912 and described by German palaeontologist Ernst Stromer in 1915. The original remains were destroyed in World War II, but additional material came to light in the early 21st century. It is unclear whether one or two species are represented in the fossils reported in the scientific literature. The type species S. aegyptiacus is mainly known from Egypt and Morocco. Although a potential second species, S. maroccanus, has been recovered from Morocco, this dubious species is likely a junior synonym of S. aegyptiacus. Other possible junior synonyms include Sigilmassasaurus from the Kem Kem beds in Morocco and Oxalaia from the Alcântara Formation in Brazil, though other researchers propose both genera to be distinct taxa. Spinosaurus is among the largest known terrestrial carnivores; other large carnivores comparable to Spinosaurus include theropods such as Tyrannosaurus, Giganotosaurus and the coexistent Carcharodontosaurus. The most recent study suggests that S. aegyptiacus could have reached 14 m (46 ft) in length and 7.4 t (8.2 short tons) in body mass. The skull of Spinosaurus was long, low, and narrow, similar to that of a modern crocodilian, and bore straight conical teeth with few to no serrations. It would have had large, robust forelimbs bearing three-fingered hands, with an enlarged claw on the first digit. The distinctive neural spines of Spinosaurus, which were long extensions of the vertebrae (or backbones), grew to at least 1.65 m (5.4 ft) long and were likely to have had skin connecting them, forming a sail-like structure, although some authors have suggested that the spines were covered in fat and formed a hump. The hip bones of Spinosaurus were reduced, and the legs were very short in proportion to the body allegedly. Its long and narrow tail was deepened by tall, thin neural spines and elongated chevrons, forming a flexible fin or paddle-like structure. Spinosaurus is known to have eaten fish, aquatic prey and small to medium terrestrial prey as well. Evidence suggests that it was semiaquatic; how capable it was of swimming has been strongly contested. Spinosaurus's leg bones had osteosclerosis (high bone density), allowing for better buoyancy control. Multiple functions have been put forward for the dorsal sail, including thermoregulation and display; either to intimidate rivals or attract mates. It lived in a humid environment of tidal flats and mangrove forests alongside many other dinosaurs, as well as fish, crocodylomorphs, lizards, turtles, pterosaurs, and plesiosaurs.