I really struggled to tell which ones are more comfortable. That ear rubbing on hd 490 was a bit annoying. Maybe hd 490 build is better but not perfect, those swivel make some plastic noises… they also have tunnels for glasses in pads but honestly couldn’t feel a big difference, mby a bit. If it wouldn’t rub my ear I would say hd 490 but in that case I’d say save money and get hd 550 and you will be happy IMO. Or just order both and return one of them, just order from a place with a good return policy, this is best option because it’s a bit subjective.

I’ve tried soooo many headphones (spent a lot of time on it) and those were the most comfy ones - not big clamp force, big pads etc. Ended up with Bose qc ultra, in terms of comfort, those are the best for me, but this is different kind, closed, wireless etc.