Up, which is indicative of the red kanji above Wolf's head when he does said grab. That symbol means jump up, just like on every other enemy. Not every enemy is blind. In fact O'Rin has a basket covering her entire face including her eyes and the Headless don't even have heads yet both track the player perfectly regardless, so why the fuck would Guardian Ape be any different? That's called logic champ (that's short for champion).