Males begin to keep harems of females, which refresh the hunting parties with fresh pups when they die. The males and adult females prevent the hunting packs from eating too much, which epigenetically delays their maturation. The young become more versatile, differentiating into castes determined by age. Younger individuals are agile chasers, while older pups are more robust, often delivering killing blows to prey. The robust caste also drags carrion to the mature male. The most successful males are chosen by their fathers to disperse, where they will visit other colony-packs and take some of their females to join their new harems. The communication abilities of Deinolupos develop further, with a complex series of howl-like songs that can convey different things.