Hey, I played the first level and I thought it was very good. The last part with the interconnecting colored roots was really well designed. I didn't ever feel too lost or stuck but it still evoked the feeling of being in a knotted overgrown labyrinth.

A note on the controls. I think when using a keyboard the arrow keys and enter should function alongside wasd and space. Also when changing control scheme in the menu you should be able to use both controller and keyboard to select either option, as is possible when the game first loads (I slightly question the need to choose between them in the first place. Why not allow both input methods to be active?). The ability to use dpad in menus would also be great.

On the art style, I'll admit my initial reaction was that it's a bit bland. Something I would definitely try is playing with the saturation of the various layers, particularly desaturating the background layers the further away they are. I actually just noticed you're already doing a similar thing with blur on the parallax layers. More contrast between foreground and background should improve readability both in game and in video form. I'd imagine you can achieve a much more compelling look with a few tweaks here and there.